A. C.  Hamel-Leigue, Herzog, S. K., Mann, D. J., Larsen, T. H., Gill, B. D., Edmonds, W. D., and Spector, S., ?Distribution and natural history of the dung beetle tribe Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) in Bolivia?, Kempffiana, vol. 5, pp. 43-95, 2009.